Velothi is another name for the Chimer who followed the prophet Veloth out from Summerset Isles to Morrowind. These were the ones who later became the Dunmer. Thus, Velothi architecture is the architecture of the early Chimer. The towers are not temples, just built in the same style. As for them not being "towers", I don't think buildings necessarily have to be [censored]-shaped to pass as towers. The strongholds were heavily influenced by Dwemer architecture, when you look at them. They're making with stones what the dwemer did with metal.
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